Leaves whorled

The following is a list of native or naturalised species which, at least some of the time, possess this spot-character.

FamilySpeciesComment
Ceratophyllaceae Ceratophyllum demersum var. demersum
Clusiaceae Garcinia livingstonei
Combretaceae Combretum adenogonium Leaves usually whorled
Combretaceae Combretum collinum Leaves sometimes whorled
Combretaceae Combretum molle Leaves rarely whorled
Combretaceae Combretum zeyheri Leaves sometimes whorled
Euphorbiaceae Croton pseudopulchellus Leaves whorled or subopposite
Lamiaceae Premna senensis
Rubiaceae Rubia horrida
Stilbaceae Nuxia congesta
Vitaceae Ampelocissus obtusata subsp. kirkiana
Number of species: 11


Copyright: Mark Hyde, Bart Wursten, Petra Ballings and Meg Coates Palgrave, 2014-25

Hyde, M.A., Wursten, B.T., Ballings, P. & Coates Palgrave, M. (2025). Flora of Botswana: List of native or naturalised species with each spot-character: Leaves whorled.
https://www.botswanaflora.com/speciesdata/species-spot-display.php?species_id=701, retrieved 28 March 2025

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